Dr. Jerri Bartholomew deciphered the complex C. shasta life cycle and identified the infectious agent for fishes in the mid-1990s.  The complex lifecycle is now maintained in the AAHL.  This myxozoan parasite (formerly Ceratomyxa shasta) causes enteronecrosis  in salmon and trout.  C. shasta is  native to the Pacific Northwest and can cause disease outbreaks in trout hatcheries and in free-ranging Chinook and coho salmon under environmental stress.  The discovery that the parasite requires a polychaete host has enabled further research on fish susceptibility, parasite genetics, and the influence of temperature and flow on infection dynamics.

Bartholomew, J., Whipple, M., Steens, D., and Fryer, J.  1997.  The life cycle of Ceratomyxa shasta, a myxosporean parasite of salmonids, requires a freshwater polychaete as an alternate host.  J. of Parasitology.